Networking / Web Browsing / E-Mail
Network setup in Ubuntu, both wired and wireless, was quite easy. My notebook's wireless adapter was detected and worked fine; all I had to do was supply my network name and I was in business. One thing that did worry me was how my unconfigured wireless card seemed to try and seek out whatever available connection there was without notifying me -- at first it attempted to connect to a neighbor's unsecured wireless base station before I redirected it back towards my own.
Both Vista and Ubuntu also let you create network profiles, although the way they're managed is markedly different. Ubuntu only lets you switch between profiles manually; Vista is semi-automatic (it makes a best guess to determine where you are), but can be manually overridden. Network connection sharing, though, is much harder to set up in Ubuntu than it is in Vista, since there's no GUI interface in Ubuntu for doing such a thing. I was able to connect to Vista's shared folders from Ubuntu, but you need to do so via a username/password combination that's valid on the Vista system you're trying to access.
Ubuntu Linux
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Windows Vista
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Web browsing is another area where the playing field is relatively level between operating systems, thanks to the general success of Firefox. Firefox is loaded as the default browser in Ubuntu, and if you don't like Internet Explorer in Vista, you can swap it out for Firefox (or most any other browser written for Windows). The behavior of Firefox on both platforms is remarkably similar; in fact, I was able to get support for Flash plugins in Ubuntu by simply pointing Firefox at a Flash-driven page and letting it download the needed components.
Ubuntu's default e-mail client is Evolution, which connects not only to POP accounts and conventional Unix mailboxes, but can also talk to Exchange servers (via Outlook Web Access) and has a built-in PIM / calendaring / appointment system. Vista's Windows Mail application is a heavily rewritten version of Outlook Express, with a stripped-down calendar/appointment application, Windows Calendar, on the side, and integration with Vista's search system. If you want more sophisticated calendaring or a full PIM, you'd need to upgrade to Outlook -- so Ubuntu has another edge here in terms of what's possible right out of the box.